I.
FOREST
I.A.8.N.b. Rounded-crowned temperate or subpolar needle-leaved
evergreen
forest
I.A.8.N.b.17. PINUS VIRGINIANA FOREST ALLIANCE
Virginia Pine Forest Alliance
I.A. Evergreen forest
Concept: This alliance includes forests dominated by Pinus
virginiana and occurring in the Piedmont from Pennsylvania south to Alabama, and
ranging west into the Appalachians, Ridge and Valley, the Cumberland Plateau,
and in scattered locales of the Interior Low Plateau. Forests in this alliance
may have admixtures of Pinus taeda, Pinus echinata, Pinus pungens, and/or
Pinus
rigida. These other species, if present, can have canopy coverage between 1 and
50%. Other associated species vary with the geographic distribution of the alliance. In many associations, a dense ericaceous shrub stratum
is typical. This alliance includes both early successional forests resulting
from natural or anthropogenic disturbance and natural forests in edaphically
extreme situations. Typically, Pinus virginiana communities are short-lived as a
forest and are more common as woodland communities [see II.C.3.N.a Pinus (rigida,
pungens, virginiana) - Quercus prinus Woodland Alliance (A.677)]. Associated
species vary with the geographic distribution of the alliance.
Comments: Appalachian pine-dominated associations need to be
revisited in relation to the ecology of shortleaf pine, Pinus echinata. Are some
stands of this type ones that historically were dominated by shortleaf pine? (MP
2002-03).
Range: Forests in this alliance are possible in the Piedmont
from Pennsylvania south to Alabama, and ranging west into the Appalachians,
Ridge and Valley, the Cumberland Plateau, and in scattered locales of the
Interior Low Plateau. The range of the alliance includes parts of Alabama,
Delaware, Georgia, Kentucky, New Jersey, North Carolina, South Carolina,
Tennessee, Maryland, Pennsylvania, West Virginia, Virginia, Ohio, and Indiana.
States/Provinces: AL GA IN KY MD NC NJ OH PA SC TN VA WV
TNC Ecoregions: 43:C, 44:C, 49:C, 50:C, 51:C, 52:C, 58:P, 59:C,
61:C
USFS Ecoregions: 221Da:CCP, 221Db:CCC, 221Ea:CC?, 221Eb:CCC,
221Ec:CCC, 221Ed:CCP, 221Ef:CCC, 221Eg:CCC, 221Ha:CCC, 221Hb:CCC, 221Hc:CCC,
221He:CCC, 221Ja:CCC, 221Jb:CCC, 221Jc:CCC, 222Da:CCC, 222Dc:CCC, 222Dd:CCC,
222Dg:CCC, 222Dj:CCC, 222Eb:CCC, 222Eg:CCC, 222Eh:CCC, 222Ej:CCC, 222El:CCC,
222En:CCC, 222Eo:CCC, 222Fc:CCC, 222Fd:CCC, 222Ff:CCC, 231Aa:CCC, 231Ab:CCC,
231Ac:CCP, 231Ad:CCC, 231Ae:CCC, 231Af:CCP, 231Ag:CCP, 231Ah:CCP, 231Ai:CCP,
231Aj:CCP, 231Ak:CCP, 231Al:CCP, 231Am:CCP, 231An:CCP, 231Ao:CCP, 231Ap:CCP,
231Bc:CCC, 231Ca:CCP, 231Cb:CCP, 231Cc:CCP, 231Cd:CCC, 231Ce:CCP, 231Cf:CCP,
231Cg:CCP, 231Da:CCC, 231Dc:CCC, M221Aa:CCC, M221Ab:CCC, M221Ac:CCC, M221Ba:CC?,
M221Bd:CCP, M221Be:CCP, M221Ca:CCP, M221Cb:CCP, M221Cc:CCC, M221Cd:CCC,
M221Ce:CCC, M221Da:CCC, M221Db:CCC, M221Dc:CCC, M221Dd:CCC
Federal Lands: DOD (Fort Jackson); NPS (Chickamauga-Chattanooga,
Great Smoky Mountains, Kennesaw Mountain, Kings Mountain, Mammoth Cave, Shiloh);
TVA (Tellico); USFS (Bankhead, Chattahoochee, Cherokee, Daniel Boone, George
Washington, Jefferson, Land Between the Lakes?, Nantahala, Oconee, Pisgah,
Sumter, Talladega, Uwharrie?)
Synonymy: IA7a. Xeric Shortleaf Pine - Oak Forest, in part
(Allard 1990); Appalachian pine-oak forest, in part (Evans 1991);
Pine--Oak/Heath, in part (Schafale and Weakley 1990); Pine--Oak/Heath, in part
(Nelson 1986); Virginia Pine CUPL, BR, RV (Pyne 1994); Virginia Pine - Mixed
Oaks HR (Pyne 1994); Virginia Pine: 79, in part (Eyre 1980)
References: Allard 1990, Andreu and Tukman 1995, Barden 1977,
Burns and Honkala 1990a, Chapman 1957, Cooper 1963, Evans 1991, Eyre 1980,
Faber-Langendoen et al. 1996, Frothingham et al. 1926, Gettman 1974, Malter
1977, Nelson 1986, Pyne 1994, Racine 1966, Schafale and Weakley 1990, Whittaker
1956
Authors: D.J. ALLARD/K.D. PATTERSO, RW, Southeast Identifier:
A.131
PINUS
VIRGINIANA - PINUS (RIGIDA, ECHINATA)
- (QUERCUS PRINUS)
/ VACCINIUM
PALLIDUM FOREST
Virginia Pine - (Pitch Pine, Shortleaf Pine) - (Rock Chestnut Oak) / Hillside
Blueberry Forest
Appalachian Low-Elevation Mixed Pine / Hillside Blueberry Forest G4? (01-02-11)
Ecological Group (SCS;MCS): Appalachian Highlands Xeric Virginia / Shortleaf
Pine Woodlands (401-27; 2.5.3.1)
Concept: This community includes
Pinus virginiana-dominated forests of low-elevation ridges and steep
upper slopes, occurring primarily in the Appalachian provinces of the eastern
United States, from central Pennsylvania, south and west to northern Georgia and
northern Alabama. This community occurs on narrow ridges, steep slopes, and
other exposed topographic positions, over shallow, infertile soils. This mainly
evergreen forest is often of low stature, with a somewhat open to closed canopy,
sparse to very dense shrub cover dominated by ericaceous species, and a sparse
herb stratum. Pinus virginiana is the canopy dominant throughout the
range of the type. In some parts of the range, other Pinus species may be
canopy associates, as well as dry site Quercus species (e.g., Quercus
prinus, Quercus coccinea). Deciduous species may form a subcanopy or sapling
stratum, particularly in areas where fire has been excluded. Common shrub
dominants include Vaccinium pallidum, Vaccinium stamineum, and Kalmia
latifolia. Herbs vary with geography but are typical of infertile, xeric
habitats. Some typical herbs in this forest are Baptisia tinctoria,
Chimaphila maculata, Dichanthelium commutatum, Epigaea repens, Euphorbia
corollata, Galax urceolata, Hypoxis hirsuta, Iris verna, Pityopsis graminifolia
var. latifolia, Pteridium aquilinum var. latiusculum, and Schizachyrium
scoparium.
Comments: Some vegetation
formerly placed (at least conceptually) in the Pinus virginiana - Quercus (coccinea,prinus)
Forest Alliance (A.408) and its provisional association Pinus virginiana
- Quercus (coccinea, prinus) Forest (CEGL005040), has been transferred here,
with this association (CEGL007119) becoming more geographically inclusive. In
Indiana examples, the substrate is primarily a matrix of acidic siltstone,
shale, and sandstone. Rarely are cliffs formed; instead the setting is mostly
very steep slopes with high hills and deep ravines. This association also
includes vegetation from the transition between the Cumberland Plateau /
Southern Ridge and Valley and the Upper East Gulf Coastal Plain in Alabama.
Though located in the Coastal Plain, these occurrences are physiographically and
floristically similar to this montane association. Early successional vegetation
associated with old fields, old pastures, clearcuts, and burned or eroded areas
and dominated by Pinus virginiana is classified as Pinus virginiana
Successional Forest (CEGL002591). Appalachian xeric oak forests with similar
floristics, but with a mainly deciduous canopy are classed in Quercus (prinus,
coccinea) / Kalmia latifolia / (Galax urceolata, Gaultheria procumbens)
Forest (CEGL006271). Appalachian shale forests and woodlands with Pinus
virginiana occur on steep, shaley slopes and have a stunted canopies and
sparse herb and shrub strata, characterized by species adapted to shaley
substrates. These shale communities are classed in Pinus virginiana - Quercus
(coccinea, prinus) Forest Alliance (A.408) and Pinus (rigida, pungens,
virginiana) - Quercus prinus Woodland Alliance (A.677).
Range: This community occurs
primarily in the Appalachian region of the United States, ranging from central
Pennsylvania, south and west through the Ridge and Valley, Blue Ridge, and
Cumberland Plateau to northern Georgia and Alabama, extending westward to
scattered areas in the Interior Low Plateau and eastward into the upper
Piedmont. It is reported from the states of Georgia, North Carolina, South
Carolina, Tennessee, Kentucky, Pennsylvania, Indiana, Ohio, and is probably in
Maryland, Virginia, and West Virginia.
States/Provinces: AL:S?, GA:S?,
IN:S?, KY:S5,S5, MD?, NC:S4, OH:S?, PA:S?, SC:S?, TN:S?, VA?, WV:S?
TNC Ecoregions: 43:C, 44:C, 49:C,
50:C, 51:C, 52:C, 59:C, 61:P
USFS Ecoregions: 221Ea:CC?,
221Eb:CCC, 221Ec:CCC, 221Ed:CCP, 221Ef:CCC, 221Eg:CCC, 221Ha:CCC, 221Hb:CCC,
221Hc:CCP, 221He:CCC, 221Ja:CCC, 221Jb:CCC, 222Da:CCC, 222Dc:CCC, 222Dg:CCC,
222Dj:CCC, 222Eg:CCC, 222Ej:CCC, 222El:CCC, 222En:CCC, 222Eo:CCC, 222Fd:CCC,
222Ff:CCC, 231Aa:CCC, 231Ab:CCC, 231Ae:CCC, 231Bc:CCC, 231Cd:CCC, 231Da:CCC,
231Dc:CCC, M221Aa:CCP, M221Ab:CCC, M221Ac:CCC, M221Bd:CCP, M221Be:CCP,
M221Cd:CCC, M221Dc:CCC, M221Dd:CCC
Federal Lands: NPS
(Chickamauga-Chattanooga, Great Smoky Mountains, Kennesaw Mountain, Kings
Mountain, Mammoth Cave); USFS (Bankhead, Chattahoochee, Cherokee, Daniel Boone,
Land Between the Lakes?, Nantahala, Pisgah, Sumter, Talladega)
Synonymy: Xeric Pine Forest, Pine
- Heath Ridge Forest (Ambrose 1990a) B, Virginia Pine - Mixed Oaks, HR (Pyne
1994) B, Virginia Pine, BR, R&V, CUPL (Pyne 1994) B, IA7c. Xeric Virginia Pine
Ridge Forest (Allard 1990) B, Virginia Pine: 79 (Eyre 1980) B, Virginia Pine -
Oak: 78 (Eyre 1980) B, Virginia pine forest (CAP pers. comm. 1998), Oligotrophic
Forest (Rawinski 1992) B, Low Mountain Pine Forest (Montane Pine Subtype)
(Schafale 1998b)
References: Allard 1990, Ambrose
1990a, Barden 1977, Burns and Honkala 1990a, CAP pers. comm. 1998, Cooper 1963,
Core 1966, Evans 1991, Eyre 1980, Fike 1999, Gettman 1974, Homoya pers. comm.,
Malter 1977, NatureServe Ecology - Southeastern U.S. unpubl. data, Nelson 1986,
Patterson et al. 1999, Peet et al. 2002, Pyne 1994, Racine 1966, Rawinski 1992,
Schafale 1998b, Schafale and Weakley 1990, Walton et al. 1997, Whittaker 1956
Authors: K.D. Patterson, SCS
Confidence: 1 Identifier: CEGL007119 - Maryland Vegetation Classification Subset Report I.A.
Evergreen forest
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